The authors thank G. Rodríguez, G. Sánchez-Montes, A. Vallvé, J. van Alphen, A. Zuiderwijk and Agentes Forestales de la Comunidad de Madrid for help in collecting individuals and P. Ochoa for technical support. Two anonymous reviewers provided valuable feedback on a previous draft. This research was funded by grants CGL2008-04271-C02-01/BOS and CGL2011-28300 (Ministerio de Ciencia e Innovación), Ministerio de Economía y Competitividad and FEDER) and PPII10-0097-4200 (Junta de Comunidades de Castilla la Mancha) to IMS. J. Gutiérrez-Rodríguez was supported by the Consejo Superior de Investigaciones Científicas of Spain (CSIC) and the European Social Fund (ESF) with a JAE-pre PhD fellowship. IMS was funded by the project ‘Biodiversity, Ecology and Global Change’, co-financed by North Portugal Regional Operational Programme 2007/2013 (ON.2–O Novo Norte), under the National Strategic Reference Framework (NSRF), through the European Regional Development Fund (ERDF) and by funding from the Spanish Severo Ochoa Program (SEV-2012-0262). IMS’s stay in Leiden was supported by a Naturalis Temminck fellowship.
Received: 12 January 2016
Revised and accepted: 15 June 2016
Published online: 14 February 2017
Editor: A. Ivanović