Contributions to Zoology, 73 (4) (2004)Andrea Grill; Rob de Vos; Jan van Arkel: The shape of endemics: Notes on male and female genitalia in the genus Maniola (Schrank, 1801), (Lepidoptera, Nymphalidae, Satyrinae)
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Maniola nurag (Figs. 3C, 4C)

Male: Gnathos substantially swollen at the base, vesica round at its extremity. Valvae considerably smaller than in M. jurtina but larger than in M. telmessia. Dorsal process clearly visible, round, flatter than in M. jurtina; distal process clearly pointed, sharper than in M. jurtina; ventral edge curved, similar to M. jurtina. Julien organ present, more fragile than in M. jurtina, but stronger than in M. telmessia.

Female: Ductus bursae similar to other Maniola species; Bursa with two elongated signa, which vary considerably in length and visibility (Fig. 5B).

FIG2

Fig. 4. Female genital apparatus of different Maniola species in comparison. (A) M. jurtina, (B) M. telmessia, (C) M. nurag, (D) M. chia,(E) M. halicarnassus, and (F) M. cypricola.