Contributions to Zoology, 73 (4) (2004)Andrea Grill; Rob de Vos; Jan van Arkel: The shape of endemics: Notes on male and female genitalia in the genus Maniola (Schrank, 1801), (Lepidoptera, Nymphalidae, Satyrinae)
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Maniola jurtina (Figs. 3A, 4A)

Male: Gnathos markedly swollen at the base, then quickly narrowing. Valvae bigger than all other species except M. halicarnassus, in shape most similar to M. nurag, but with a characteristic curve towards the distal process; distal and dorsal process round; ventral edge different from the other Maniola species, most similar to M. nurag. Julien organ always clearly visible, very thick and strong, can be twice the size as in congenerics.

FIG2

Fig. 3. Male genital apparatus of different Maniola species. (A) M. jurtina, (B) M. telmessia, (C) M. nurag, (D) M. chia,(E) M. halicarnassus, (F) M. cypricola, and (G) intermediate form (from Sardinia).

Female: Length of ductus bursae comparable to the other species; notably in none of the dissected females we found signa, although they were clearly visible in all female individuals of the other species. The absence of these marks might be a good distinctive characteristic between M. jurtina and the other species in the genus Maniola.